

Laetoli Giant Tortoise
Geochelone laetoliensis
🐢🌍
Chordata
Reptilia
Testudines
Crytodira
Testudinidae
Aldabrachelys
Geochelone laetoliensis
Geochelone = “earth tortoise” (Greek); laetoliensis = “from Laetoli.”
A robust Pliocene tortoise from Laetoli, living alongside early hominins.
Description
Geochelone laetoliensis was a large land tortoise inhabiting the Pliocene savanna ecosystems of northern Tanzania around 3.5 million years ago. Its anatomy resembled modern giant tortoises—broad domed shell, thick limb bones, and a slow but powerful terrestrial gait. Fossils suggest a heavy, high-domed carapace suited to protection against predators in open habitats.
Like other giant Geochelone, it likely lived a slow-paced, herbivorous lifestyle, grazing on grasses, forbs, and succulent plants. Its size may have exceeded that of most modern mainland tortoises, making it one of the larger terrestrial reptiles of its ecosystem.
“Aldabrachelys” laetoliensis (Meylan and Auffenberg 1987)
(Late Pliocene to Early Pleistocene)
Laetoli Giant Tortoise Tanzania
Size: CL ca. 100 cm
Geochelone (Aldabrachelys) laetoliensis † Meylan and Auffenberg 1987:1167 [Late Pliocene, Laetoli Beds, Olduvai Gorge, Tanzania], Geochelone laetoliensis, Aldabrachelys laetoliensis, Stigmochelys laetoliensis
Comment: This species was recorded from hominin habitation sites in Olduvai from about 3,800,000–2,600,000 ybp and was exposed to early human exploitation (Harrison 2011). This species has tentatively been assigned to both Aldabrachelys and Stigmochelys (to which it may be closer; F. Lapparent de Broin, pers. comm.), but further material and new analysis may well indicate that it
belongs to a new genus
Quick Facts
Max Mass
Shoulder Height
Standing Height
Length
Diet
Trophic Level
40
1
kg
m
m
m
Herbivore Generalist
Grazer
Hunt History
This species comes from the famed Laetoli fossil beds (~3.6–3.0 Ma), known for early hominin footprints. While G. laetoliensis does not show direct evidence of human hunting—modern humans were far in the future—its remains appear in the same deposits documenting early Australopithecus afarensis. Its presence helps reconstruct the paleoenvironment as a warm, semi-arid mosaic of grassland and shrubland.
Three examples:
• Found in Laetoli Beds (Tanzania), the same locale as the iconic Australopithecus foot tracks.
• Pliocene fauna association includes early suids, equids, and bovids—indicating open savanna.
• Represents the typical large testudinid lineage present throughout African Plio-Pleistocene sites.
Time & Range
Extinction Status
Globally Extinct
Extinction Date
Temporal Range
Region
3000000
BP
Pliocene Epoch (5.3–2.58 Ma)
East Africa (Laetoli, Tanzania)
Fat Analysis
Fatness Profile:
Low
Fat %
7
Est. Renderable Fat
3
kg
Targeted Organs
Adipose Depots
Preferred Cuts
Hunt Difficulty (x/5)
1





